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After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

(FOX 5/KUSI) — Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died. The zoo said Gramma had ongoing bone conditions and that caretakers had to make the ...

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White House. By Victor Mather Tortoises are slow. While most of Earth’s creatures zip ...

This is read by an automated voice. Please report any issues or inconsistencies here. Gramma, a beloved Galápagos tortoise at the San Diego Zoo believed to be 141 years old, was euthanized after ...